During World War II, U.S. Navy submarines paid heavily for their success. A total of 374 officers and 3131 men remain on board 52 U.S. submarines still on patrol: Albacore, Amberjack, Argonaut, Barbel, Bonefish, Bullhead, Capelin, Cisco, Corvina, Darter, Dorado, Escolar, Flier, Golet, Grampus, Grayback, Grayling, Grenadier, Growler, Grunion, Gudgeon, Harder, Herring, Kete, Lagarto, Perch, Pickerel, Pompano, Robalo, Runner, R-12, Scamp, Scorpion, Sculpin, Sealion, Seawolf, Shark I, Shark II, Snook, Swordfish, S-26, S-27, S-28, S-36, S-39, S-44, Tang, Trigger, Triton, Trout, Tullibee, and Wahoo. Fleet Admiral C.W. Nimitz said that submarines held the lines against the enemy while fleets replaced losses and repaired wounds, and Vice Admiral C.A. Lockwood, Jr., said they went down fighting and that their surviving brothers took a grim toll on the enemy to avenge their deaths.