During World War II, U.S. Navy submarines paid heavily for their success, with 374 officers and 3131 men aboard 52 U.S. submarines still on patrol: Albacore, Amberjack, Argonaut, Barbel, Bonefish, Bullhead, Capelin, Cisco, Corvina, Darter, Dorado, Escolar, Flier, Golet, Grampus, Grayback, Grayling, Grenadier, Growler, Grunion, Gudgeon, Harder, Herring, Kete, Lagarto, Perch, Pickerel, Pompano, Robalo, Runner, R-12, Scamp, Scorpion, Sculpin, Sealion, Seawolf, Shark I, Shark II, Snook, Swordfish, S-26, S-27, S-28, S-36, S-39, S-44, Tang, Tiger, Triton, Trout, Tullibee, and Wahoo. Fleet Admiral C.W. Nimitz said that submarines held the lines against the enemy while fleets replaced losses and repaired wounds, and Vice Admiral C.A. Lockwood, Jr., said they went down fighting and that the brothers who survived them took a grim toll on their enemy to avenge their deaths.